Ecology Summary - Cottus asper

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Main Ref.	Brown, L.R., S.A. Matern and P.B. Moyle, 1995
Remarks	There appears to be two genetically distinct forms (Ref. 27547), an inland form found in sandy and rocky shores of lakes, and a coastal form usually found over sand in quiet runs of small to medium rivers; sometimes in salt water near river mouths (Ref. 2850). The coastal form moves into brackish estuaries to spawn (Ref. 27547). Feed mainly on aquatic insect larvae and bottom invertebrates (Ref. 1998). Juvenile and adults feed mainly on insects (Ref. 13348).

Main Ref.

Brown, L.R., S.A. Matern and P.B. Moyle, 1995

Remarks

There appears to be two genetically distinct forms (Ref. 27547), an inland form found in sandy and rocky shores of lakes, and a coastal form usually found over sand in quiet runs of small to medium rivers; sometimes in salt water near river mouths (Ref. 2850). The coastal form moves into brackish estuaries to spawn (Ref. 27547). Feed mainly on aquatic insect larvae and bottom invertebrates (Ref. 1998). Juvenile and adults feed mainly on insects (Ref. 13348).

Aquatic zones / Water bodies

	Marine - Neritic	Marine - Oceanic	Brackishwater	Freshwater
Marine zones / Brackish and freshwater bodies	supra-littoral zone littoral zone sublittoral zone	epipelagic mesopelagic epipelagic abyssopelagic hadopelagic	estuaries/lagoons/brackish seas mangroves marshes/swamps	rivers/streams lakes/ponds caves exclusively in caves

Highighted items on the list are where Cottus asper may be found.

Habitat

Substrate	Soft Bottom: sand; Hard Bottom: rocky;
Substrate Ref.	Morrow, J.E., 1980
Special habitats
Special habitats Ref.

Associations

Ref.
Associations
Associated with
Association remarks
Parasitism

Feeding

Feeding type	mainly animals (troph. 2.8 and up)
Feeding type Ref.	Brown, L.R., S.A. Matern and P.B. Moyle, 1995
Feeding habit	hunting macrofauna (predator)
Feeding habit Ref.	Brown, L.R., S.A. Matern and P.B. Moyle, 1995

Trophic Level(s)

Estimation method	Original sample		Unfished population		Remark
Estimation method	Troph	s.e.	Troph	s.e.	Remark
From diet composition	3.20		3.10	0.40	Troph of adults from 1 study.
From individual food items	3.61	0.51			Trophic level estimated from a number of food items using a randomized resampling routine.
Ref.	Brown, L.R., S.A. Matern and P.B. Moyle, 1995

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