Africa: Lake Tanganyika and associated rivers (Ref. 4983), including the Lukuga (up to Kisimba-Kilia rapids) connecting the lake to the upper Congo River (Ref. 53528, 93587), the Malagarazi (Ref. 54847) and the Rusizi (Ref. 87863). Also in Akagera system, where it might have been introduced (Ref. 46152).