Metamorphosed females distinguished by the following characteristics: escal bulb with distal prolongation, conical at base and distally more or less deeply bifurcated; short posterior appendage, conical or compressed; absence of additional escal appendage; barbel with long undivided stem and a distal tassel of branches; stem of barbel without filaments or lateral branches; distal tassel more or less distinctly divided into 3-4 basal and 3 terminal branches; juvenile female with concentration of subdermal pigment on caudal peduncle; parasitic males with pointed sphenotic spines (Ref. 86949).