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	<dc:identifier>FB-727</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=727</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Elasmobranchii</dwc:Class>
	<dwc:Order>Squatiniformes</dwc:Order>
	<dwc:Family>Squatinidae</dwc:Family>
	<dwc:Genus>Squatina</dwc:Genus>
	<dwc:ScientificName>Squatina argentina (Marini, 1930)</dwc:ScientificName>
	<commonName xml:lang='pt'>Anjo africano</commonName>
	<commonName xml:lang='pt'>Anjo-do-leste-africano</commonName>
	<commonName xml:lang='ar'>قِرش مَلَك إفريقي</commonName>
	<commonName xml:lang='da'>Afrikansk havengel</commonName>
	<commonName xml:lang='de'>Afrikanischer Engelhai</commonName>
	<commonName xml:lang='en'>African angel shark</commonName>
	<commonName xml:lang='en'>African angelshark</commonName>
	<commonName xml:lang='et'>Aafrika ingelhai</commonName>
	<commonName xml:lang='fr'>Ange de mer africain</commonName>
	<commonName xml:lang='es'>Angelote africano</commonName>
	<commonName xml:lang=''>Afrikaanse zee-engel</commonName>
	<commonName xml:lang=''>Violon</commonName>
	<commonName xml:lang=''>非洲扁鯊</commonName>
	<commonName xml:lang=''>非洲扁鲨</commonName>
	<dcterms:created>1992-06-15</dcterms:created>
	<dcterms:modified>2007-02-12</dcterms:modified>
	<reference isbn='92-5-103623-3.' url='http://www.fishbase.org/references/FBRefSummary.php?id=171&amp;speccode=726'>Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter0 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p. (Ref. 171)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=726'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=247&amp;speccode=726'>Compagno, L.J.V.0 FAO Species Catalogue. Vol. 4. Sharks of the world. An annotated and illustrated catalogue of shark species known to date. Part 1 - Hexanchiformes to Lamniformes. FAO Fish. Synop. 125(4/1):1-249. Rome, FAO. (Ref. 247)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=726'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5213&amp;speccode=726'>Fischer, W., I. Sousa, C. Silva, A. de Freitas, J.M. Poutiers, W. Schneider, T.C. Borges, J.P. Feral and A. Massinga0 Fichas FAO de identificaçao de espécies para actividades de pesca. Guia de campo das espécies comerciais marinhas e de águas salobras de Moçambique. Publicaçao preparada em collaboraçao com o Instituto de Investigaçao Pesquiera de Moçambique, com financiamento do Projecto PNUD/FAO MOZ/86/030 e de NORAD. Roma, FAO. 1990. 424 p. (Ref. 5213)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=726'>Compagno, L.J.V., D.A. Ebert and M.J. Smale0 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6588&amp;speccode=726'>Bass, A.J.0 Squatinidae. p. 107. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 6588)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10311&amp;speccode=726'>Fritzsch, B. and P. Moller0 A history of electroreception. p. 39-55. In P. Moller (ed.) Electric fishes:history and behavior. Fish and Fisheries Series 17. Chapman &amp; Hall, London. (Ref. 10311)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=11228&amp;speccode=726'>Heemstra, P.C.0 Additions and corrections for the 1995 impression. p. v-xv. In M.M. Smith and P.C. Heemstra (eds.) Revised Edition of Smiths' Sea Fishes. Springer-Verlag, Berlin. (Ref. 11228)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12484&amp;speccode=726'>van der Elst, R.0 A guide to the common sea fishes of southern Africa. (3rd Ed.). Struik Publishers, Cape Town. 398 p. (Ref. 12484)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26388&amp;speccode=726'>Sanches, J.G.0 Nomenclatura Portuguesa de organismos aquáticos (proposta para normalização estatística). Publ. Avuls. Inst. Nac. Invest. Pescas 14:322 p. (Ref. 26388)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=726'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=726'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='3-87429-411-0' url='http://www.fishbase.org/references/FBRefSummary.php?id=33390&amp;speccode=726'>Fricke, R.0 Fishes of the Mascarene Islands (Réunion, Mauritius, Rodriguez): an annotated checklist, with descriptions of new species. Koeltz Scientific Books, Koenigstein, Theses Zoologicae, Vol. 31:759 p. (Ref. 33390)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=33559&amp;speccode=726'>Cortés, E.0 Standardized diet compositions and trophic levels of sharks. ICES J. Mar. Sci. 56:707-717. (Ref. 33559)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=35509&amp;speccode=726'>Anonymous0 Fish collection database of the Intitut fur Seefischerei (ISH). BIOLAB Forschungsinstitut, Hohenwestedt, Germany. (Ref. 35509)</reference>
	<reference isbn='0-8018-6048-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=35766&amp;speccode=726'>Compagno, L.J.V.0 Checklist of living elasmobranchs. p. 471-498. In W.C. Hamlett (ed.) Sharks, skates, and rays: the biology of elasmobranch fishes. Johns Hopkins University Press, Maryland. (Ref. 35766)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36670&amp;speccode=726'>Anonymous0 Fish collection database of the J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. (Ref. 36670)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=43454&amp;speccode=726'>Bor, P.H.F.0 Nederlandse naamlijst van de recente haaien en roggen (Chondrichthyes: Elasmobranchii) van de wereld. World Wide Web electronic publication www.rajidae.tmfweb.nl, version (05/2002). (Ref. 43454)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50449&amp;speccode=726'>Dulvy, N.K. and J.D. Reynolds0 Evolutionary transitions among egg-laying, live-bearing and maternal inputs in sharks and rays. Proc. R. Soc. Lond., Ser. B: Biol. Sci. 264:1309-1315. (Ref. 50449)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51471&amp;speccode=726'>Carl, H.0 Danish fish names. Zoological Museum of Copenhagen. Unpublished. (Ref. 51471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58078&amp;speccode=726'>Baissac, J. de B.0 SWIOP/WP/54 - Checklist of the marine fishes of Mauritius. RAF/87/008/WP/54/90 Regional Project for the Development &amp; Management of Fisheries in the Southwest Indian Ocean. (Ref. 58078)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=726'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=75549&amp;speccode=726'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis, 29 April 2008. (Ref. 75549)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=726'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=726'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=726'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=97038&amp;speccode=726'>Aura, C.M., C.N. Munga, E. Kimani, J.O. Manyala and S. Musa0 Length- weight relationships for nine deep sea fish species off the Kenyan coast. Pan-American Journal of Aquatic Sciences 6(2):188-192. (Ref. 97038)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=726'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=726'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=726'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118272&amp;speccode=726'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=128961&amp;speccode=726'>Weigmann, S., D.F.B. Vaz, K.V. Akhilesh, R.H. Leeney and G.J.P. Naylor0 Revision of the western Indian ocean angel sharks, genus &lt;i&gt;Squatina&lt;/i&gt; (Squatiniformes, Squatinidae), with description of a new species and redescription of the African angel shark &lt;i&gt;Squatina africana&lt;/i&gt; Regan, 1908. Biology 2023, 12(77):975. (Ref. 128961)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=132125&amp;speccode=726'>Ferragut-Perello, F., S. Ramirez-Amaro, A.C. Tsikliras, N. Petit-Marty, D. Dimarchopoulou0 Exploitation and conservation status of the thornback ray (&lt;i&gt;Raja clavata&lt;/i&gt;) in the Balearic Islands (Western Mediterranean). Fishes 8(2):117. (Ref. 132125)</reference><dataObject>
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	<dcterms:created>1995-05-17</dcterms:created>
	<dcterms:modified>2024-08-05</dcterms:modified>
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	<dc:rights>This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged</dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=726</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology</subject>
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	<agent homepage='' logoURL='' role='recorder'>Kent E. Carpenter</agent>
	<dcterms:created>1992-06-15 00:00:00</dcterms:created>
	<dcterms:modified>2023-07-17 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<dc:description>Found on the continental shelf and uppermost slope, from the surf line close inshore to 494 m depth (Ref. 247).  Feeds on bony fish, squid and octopus (Ref. 247).  Ovoviviparous (Ref. 50449), with about 7 to 11 young in a litter (Ref. 247).  Largest juvenile males measured with clearly immature claspers are 45.3 cm TL (ZMH 26100) and 50.3 cm TL (MNHN-IC-1987-1265) while the largest female examined is 94.0 cm TL (SAIAB 187381) and possibly an adult; while small female specimens of 29.3 cm TL (ZMMU P-14844), 30.9 cm TL (ZMH 123064), and 38.5 cm TL (MNHN-IC-1988-0359) mm TL are clearly juvenile (Ref. 128961). Sometimes caught by surf anglers (Ref. 6588).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=128961&amp;speccode=726'>Weigmann, S., D.F.B. Vaz, K.V. Akhilesh, R.H. Leeney and G.J.P. Naylor 2023 Revision of the western Indian ocean angel sharks, genus &lt;i&gt;Squatina&lt;/i&gt; (Squatiniformes, Squatinidae), with description of a new species and redescription of the African angel shark &lt;i&gt;Squatina africana&lt;/i&gt; Regan, 1908. Biology 2023, 12(77):975. (Ref. 128961)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>2023-07-17 00:00:00</dcterms:created>
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	<dc:description>This medium-sized angel shark species (maximum size 122.0 or possibly 130.0 cm TL) posess the following characters: no median row of scute-like denticles on the trunk; anterior nasal flap has two lateral, elongate barbels and a medial rectangular barbel, the ventral margin of medial barbel is slightly fringed, but that of lateral barbels are almost smooth; posterior nasal flap without additional barblet; concave between eyes; pectoral-pelvic space 6.0-10.2% TL; pectoral-fin apex is angular; pelvic-fin free rear tips do not reach  level of first dorsal-fin origin; tail moderately long, its length from cloaca 51.2-53.2% TL; pectoral fins are  moderately long, length 31.8-35.6% TL; dorsal fins are not lobe-like; first dorsal-fin base is somewhat longer than second dorsal-fin base; caudal fin of adults with angular apices; monospondylous centra 46-49; diplosondylous precaudal centra 58-62; total precaudal centra 104-111; total vertebral centra 134-143; pectoral-fin skeleton is with propterygium articulating with three radials. Colouration: dorsal medium to dark brown, reddish-brown or grayish, with a variable pattern of numerous light and dark reddish spots and blotches, marbled with brownish reticulations and partially forming symmetrical dark bands or saddles (Ref. 128961).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=128961&amp;speccode=726'>Weigmann, S., D.F.B. Vaz, K.V. Akhilesh, R.H. Leeney and G.J.P. Naylor 2023 Revision of the western Indian ocean angel sharks, genus &lt;i&gt;Squatina&lt;/i&gt; (Squatiniformes, Squatinidae), with description of a new species and redescription of the African angel shark &lt;i&gt;Squatina africana&lt;/i&gt; Regan, 1908. Biology 2023, 12(77):975. (Ref. 128961)</reference></dataObject><dataObject>
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	<dc:description>Western Indian Ocean:  tropical and warm temperate waters from Tanzania to the eastern Cape coast of South Africa.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=247&amp;speccode=726'>Compagno, L.J.V. 1984 FAO Species Catalogue. Vol. 4. Sharks of the world. An annotated and illustrated catalogue of shark species known to date. Part 1 - Hexanchiformes to Lamniformes. FAO Fish. Synop. 125(4/1):1-249. Rome, FAO. (Ref. 247)</reference></dataObject><dataObject>
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	<dc:description>fisheries: subsistence fisheries; gamefish: yes</dc:description>
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	<reference isbn='92-5-103623-3.' url='http://www.fishbase.org/references/FBRefSummary.php?id=171&amp;speccode=726'>Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter 1994 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p. (Ref. 171)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6588&amp;speccode=726'>Bass, A.J. 1986 Squatinidae. p. 107. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 6588)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
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	<dc:description>Ovoviviparous, with about 7 to 11 young in a litter. Size at birth between 28 and 34 cm.  Embryos feed solely on yolk (Ref. 50449).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=726'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-726</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Kent E. Carpenter</agent>
	<dcterms:created>1992-06-15 00:00:00</dcterms:created>
	<dcterms:modified>2023-07-17 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=726</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>84.0 cm TL (male/unsexed; (Ref. 128961)); 108 cm TL (female)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=128961&amp;speccode=726'>Weigmann, S., D.F.B. Vaz, K.V. Akhilesh, R.H. Leeney and G.J.P. Naylor 2023 Revision of the western Indian ocean angel sharks, genus &lt;i&gt;Squatina&lt;/i&gt; (Squatiniformes, Squatinidae), with description of a new species and redescription of the African angel shark &lt;i&gt;Squatina africana&lt;/i&gt; Regan, 1908. Biology 2023, 12(77):975. (Ref. 128961)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
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	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>John E. Randall</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject></subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-726</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1992-06-15 00:00:00</dcterms:created>
	<dcterms:modified>2023-07-17 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>John E. Randall</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=726</dc:source>
	<subject></subject>
	<dc:description>demersal; marine; depth range ? - 494 m (Ref. 5578), usually 60 - 300 m (Ref. 5578)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=726'>Compagno, L.J.V., D.A. Ebert and M.J. Smale 1989 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=726'>Compagno, L.J.V., D.A. Ebert and M.J. Smale 1989 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference></dataObject>
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