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	<dc:identifier>FB-8757</dc:identifier>
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	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Cichliformes</dwc:Order>
	<dwc:Family>Cichlidae</dwc:Family>
	<dwc:Genus>Neolamprologus</dwc:Genus>
	<dwc:ScientificName>Neolamprologus tetracanthus (Boulenger, 1899)</dwc:ScientificName>
	<commonName xml:lang='en'>Fourspine cichlid</commonName>
	<commonName xml:lang='fi'>Piikkiluola-ahven</commonName>
	<commonName xml:lang=''>四棘新亮丽鲷</commonName>
	<commonName xml:lang=''>四棘新亮麗鯛</commonName>
	<synonym relationship='synonym'>Lamprologus brevianalis Boulenger, 1906</synonym>
	<synonym relationship='synonym'>Lamprologus marginatus Boulenger, 1914</synonym>
	<synonym relationship='synonym'>Lamprologus tetracanthus Boulenger, 1899</synonym>
	<dcterms:created>1994-02-02</dcterms:created>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1672&amp;speccode=8757'>Riehl, R. and H.A. Baensch0 Aquarien Atlas. Band. 1. Melle: Mergus, Verlag für Natur-und Heimtierkunde, Germany. 992 p. (Ref. 1672)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=8757'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='0-913235-53-9' url='http://www.fishbase.org/references/FBRefSummary.php?id=4537&amp;speccode=8757'>Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea and W.B. Scott0 World fishes important to North Americans. Exclusive of species from the continental waters of the United States and Canada. Am. Fish. Soc. Spec. Publ. (21):243 p. (Ref. 4537)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5667&amp;speccode=8757'>Maréchal, C. and M. Poll0 Neolamprologus. p. 274-294. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5667)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7343&amp;speccode=8757'>Konings, A.0 Tanganyika cichlids. Verduijn Cichlids &amp; Lake Fish Movies, Zevenhuizen, Holland. 272 p. (Ref. 7343)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12818&amp;speccode=8757'>Anonymous0 Fish registrations within the museum database of the Vertebrate Section of the Royal Museum for Central Africa. MRAC, Tervuren, Belgium. (Ref. 12818)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13371&amp;speccode=8757'>Riehl, R. and H.A. Baensch0 Aquarien Atlas, Band 1. 10th edition. Mergus Verlag GmBH, Melle, Germany. 992 p. (Ref. 13371)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31402&amp;speccode=8757'>Swedish Museum of Natural History0 Fish collection database of the Naturhistoriska riksmuseet (Swedish Museum of Natural History). Ichthyology Section, Department of Vertebrate Zoology, Swedish Museum of Natural History, Stockholm, Sweden. (Ref. 31402)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=8757'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=8757'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=35508&amp;speccode=8757'>Anonymous0 The icthyological collection of the Zoological Museum Hamburg (ZMH). Division of Ichthyology and Herpetology, Zoological Museum Hamburg (ZMH). (Ref. 35508)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36670&amp;speccode=8757'>Anonymous0 Fish collection database of the J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. (Ref. 36670)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38732&amp;speccode=8757'>Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=40115&amp;speccode=8757'>Kohda, M., Y. Yanagisawa, T. Sato, K. Nakaya, Y. Niimura, K. Matsumoto and H. Ochi0 Geographical colour variation in cichlid fishes at the southern end of Lake Tanganyika. Environ. Biol. Fishes 45(3):237-248. (Ref. 40115)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41414&amp;speccode=8757'>Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51086&amp;speccode=8757'>Varjo, M., L. Koli and H. Dahlström0 Kalannimiluettelo (versio 10/03). Suomen Biologian Seura Vanamo Ry. (Ref. 51086)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=8757'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=117407&amp;speccode=8757'>Tsuboi, M., A. Husby, A. Kotrschal, A. Hayward, S.D. Büchel, J. Zidar, H. Løvlie and N. Kolm0 Comparative support for the expensive tissue hypothesis: big brains are correlated with smaller gut and greater parental investment in Lake Tanganyika cichlids. Evolution 69(1):190-200. (Ref. 117407)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118184&amp;speccode=8757'>Reddon, A.R., C.M. O'Connor, I.Y. Ligocki, J.K. Hellmann, S.E. Marsh-Rollo, I.M. Hamilton and S. Balshine0 No evidence for larger brains in cooperatively breeding cichlid fishes. Can. J. Zool. 94:373-378. (Ref. 118184)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118251&amp;speccode=8757'>Gonzalez-Voyer, A., S. Winberg and N. Kolm0 Distinct evolutionary patterns of brain and body size during adaptive radiation. Evolution 63:2266-2274. (Ref. 118251)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118717&amp;speccode=8757'>Tsuboi, M., W. van der Bijl, B.T. Kopperud, J. Erritzoe, K.L. Voje, A. Kotrschal, K.E. Yopak, S.P. Collin, A.N. Iwaniuk and N. Kolm0 Breakdown of brain-body allometry and the encephalization of birds and mammals. Nat. Ecol. Evol. 2:1495-1500. (Ref. 118717)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130435&amp;speccode=8757'>IUCN0 The IUCN Red List of Threatened Species. 1964 - onwards. https://www.iucnredlist.org/ (Ref. 130435)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137577&amp;speccode=8757'>Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577)</reference><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-02-02 00:00:00</dcterms:created>
	<dcterms:modified>2018-06-13 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Sand-dweller.  Feeds on mollusks, insect larvae, and fish.  N. tetracanthus is not a typical crusher of mollusk shells but it is able to suck the mollusk out of its shell (Ref. 7343).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5667&amp;speccode=8757'>Maréchal, C. and M. Poll 1991 Neolamprologus. p. 274-294. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5667)</reference></dataObject><dataObject>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Africa:  Endemic to Lake Tanganyika where it is widely distributed.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5667&amp;speccode=8757'>Maréchal, C. and M. Poll 1991 Neolamprologus. p. 274-294. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5667)</reference></dataObject><dataObject>
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	<dc:description>fisheries: commercial; aquarium: commercial</dc:description>
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	<reference isbn='0-913235-53-9' url='http://www.fishbase.org/references/FBRefSummary.php?id=4537&amp;speccode=8757'>Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea and W.B. Scott 1991 World fishes important to North Americans. Exclusive of species from the continental waters of the United States and Canada. Am. Fish. Soc. Spec. Publ. (21):243 p. (Ref. 4537)</reference>
	<reference isbn='0-913235-53-9' url='http://www.fishbase.org/references/FBRefSummary.php?id=4537&amp;speccode=8757'>Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea and W.B. Scott 1991 World fishes important to North Americans. Exclusive of species from the continental waters of the United States and Canada. Am. Fish. Soc. Spec. Publ. (21):243 p. (Ref. 4537)</reference></dataObject><dataObject>
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	<dc:source>tetracanthus</dc:source>
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	<dc:description>Sand-dweller. Feeds on molluscs, insect larvae, and fish. N. tetracanthus is not a typical crusher of mollusc shells but it is able to suck the mollusc out of its shell.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7343&amp;speccode=8757'>Konings, A. 1988 Tanganyika cichlids. Verduijn Cichlids &amp; Lake Fish Movies, Zevenhuizen, Holland. 272 p. (Ref. 7343)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-8757</dc:identifier>
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	<dc:title>Max. size</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>20.0 cm TL (male/unsexed; (Ref. 5667))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5667&amp;speccode=8757'>Maréchal, C. and M. Poll 1991 Neolamprologus. p. 274-294. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5667)</reference></dataObject><dataObject>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>benthopelagic; freshwater; pH range: 7.5 - 8.0; dH range: 12 - 15, usually 6 - 12 m (Ref. 118184)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118184&amp;speccode=8757'>Reddon, A.R., C.M. O'Connor, I.Y. Ligocki, J.K. Hellmann, S.E. Marsh-Rollo, I.M. Hamilton and S. Balshine 2016 No evidence for larger brains in cooperatively breeding cichlid fishes. Can. J. Zool. 94:373-378. (Ref. 118184)</reference></dataObject>
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