Environment: milieu / climate zone / depth range / distribution range
البيئة
; المياه العذبة; مياه مخلوطة قاعية التغذية و المعيشة; أسماك صاعدة (Ref. 51243). Subtropical; 10°C - 20°C (Ref. 2059); 64°N - 35°N, 12°E - 53°E
Eurasia: Caspian, Black, Marmara and Baltic Sea basins (Sweden and Finland north to 63°N), North Sea basin from Elbe to Ems drainages. In Anatolia: in Marmara basin, south to Great Menderes and Lake Egridir and east to Kızılırmak drainage. Absent between Kızılırmak and Çoruh drainages. Extirpated in Crimea. Introduced in Rhine.
الحجم / وزن / العمر
Maturity: Lm ? range ? - ? cm
Max length : 50.0 cm TL ذكر/ مختلط الجنس; (Ref. 556); common length : 20.0 cm TL ذكر/ مختلط الجنس; (Ref. 556); أعلا وزن تم نشرة: 1.4 kg (Ref. 35541); العمر: 15 سنين (Ref. 59043)
الأشواك الظهرية (المجموع) : 3; الأشعة الظهرية الناعمة (المجموع) : 8 - 9; شوكة شرجية: 3; أشعه شرجية لينه: 16 - 22. Diagnosed from its congeners in Europe by the following characters: anal fin with 16-21½ branched rays; back keeled behind dorsal base; in spawning season, males blackish brown with orange cheek and sometimes belly (Ref. 59043). Caudal fin with 19 soft rays. Scalar formula: 49-64, 6-10 (Ref. 40476).
Inhabits brackish estuaries, large to medium rivers and some large subalpine lakes. Sedentary populations inhabit even in small rivers or barbel zone. Feeds mainly on small molluscs and insect larvae. Breeds in riffles in shallow, fast-flowing streams and rivers on gravel. Semi-anadromous populations forage in freshened parts of sea and migrate for long distances to spawn. Lacustrine populations move to fast-flowing tributaries. Spent adults return to foraging habitats (Ref. 59043). Threatened from its range due to water obstruction (Ref. 26100).
Produces 15,000-20,000 eggs at the age of 6-7 years (Ref. 12259). Spawns for several years. Semi-anadromous populations undertake long distance spawning migrations while lacustrine populations migrate to fast-flowing tributaries. In Dniepr and Kuban, migration starts in September, decreases in winter and resumes in March-May (Ref. 59043).
Kottelat, M. and J. Freyhof, 2007. Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)
IUCN Red List Status (Ref. 130435)
استخدامات بشرية
مصائد: غير مهمة تجارياً; الأستزراع المائي: تجاري; لعبة سمكه: نعم
أدوات
تقارير خاصة
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مصادر علي الأنترنت
Estimates based on models
Phylogenetic diversity index (Ref.
82804): PD
50 = 0.5625 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00724 (0.00660 - 0.00795), b=3.14 (3.11 - 3.17), in cm total length, based on LWR estimates for this species (Ref.
93245).
مستوى غذائي (Ref.
69278): 3.3 ±0.1 se; based on diet studies.
المرونه (Ref.
120179): منخفض, الحد الزمني الأدني لتضاعف عدد أفراد المجتمع 4.5-14 سنة (K=0.22; tm=3; tmax=15).
Fishing Vulnerability (Ref.
59153): Moderate vulnerability (37 of 100).
